Code: Select all
\titlespacing{\section}{0pt}{*0}{*-2}
\titlespacing{\subsection}{0pt}{*1}{*-2}
\titlespacing{\subsubsection}{0pt}{*1}{*-2}
\titlespacing{\bodymath}{0pt}{*-12}{*-12}
Also the materials and methods title shouldn't have all that space above it (the top margin is ~2.5cm and looks ok on other pages).
Thanks